Grizzly 399, a beloved mama bear known as the queen of Wyoming's Grand Teton National Park, has died after being hit by a car, the U.S. Fish & Wildlife Service reports.

Grizzly 399’s ashes were spread in the Pilgrim Creek area inside Grand Teton National Park. That was her home turf and probable birth place. She died at the age of 28, having raised 18 cubs.

Grizzly 399: Mourning the mama bear who was queen of Wyoming's Grand Teton National Park The bear was a mother to at least 18 cubs over the years and the star of a PBS documentary.

Grizzly 399, seen with her cubs in June 2020, was struck and killed by a vehicle Tuesday night near Grand Teton National Park in Wyoming, the U.S. Fish and Wildlife Service announced Wednesday.

Grizzly No. 399, who was a famous female grizzly bear, was fatally struck by a vehicle in Grand Teton National Park in western Wyoming on Tuesday, Oct. 22.

In this June 2011, file photo, grizzly bear No. 399 crosses a road in Grand Teton National Park, Wyo., with her three cubs. Close Ad To stream 9NEWS on your phone, you need the 9NEWS app.

Famed grizzly 399 cremated, ashes spread in Grand Teton National Park The 28-year-old bear was killed in an accidental vehicle collision on Oct. 22, about 40 miles south of the park. At the time ...

In this undated photo provided by Grand Teton National Park a grizzly bear known as No. 399 stands along side a cub. (C. Adams/Grand Teton National Park via AP) ...

Grizzly 399, seen with her cubs in June 2020, was struck and killed by a vehicle Tuesday night near Grand Teton National Park in Wyoming, the U.S. Fish and Wildlife Service announced Wednesday.

“399 will always be part of this special place,” said Grand Teton National Park Superintendent Chip Jenkins. “However, there is still work to do to ensure her descendants and all grizzly ...

In October, grizzly 399 was struck and killed by a vehicle south of Grand Teton National Park. Law enforcement announced the driver was not speeding and the bear's death was an accident.

GRAND TETON NATIONAL PARK — The U.S. Fish and Wildlife Service and Grand Teton National Park honored the legacy of grizzly bear 399 by returning her ashes to the Pilgrim Creek area of the park.